Through a series of flashbacks, his parents are revealed to be named Lyla Novacek, (Keri Russell), a famous concert cellist, and Louis Connelly (Jonathan Rhys Meyers), an Irish guitarist and lead singer of a rock band, who spent one romantic night together and never saw each other again. Lyla became pregnant, which her father did not approve of, instead wanting Lyla to have a successful career without the obstacle of a child. After an argument with her father, Lyla ran out of a restaurant and was hit by a car. While in the hospital, she gave birth to a son. Afterwards, she was told by her father that her child had died, unaware that he had actually been delivered successfully and then been sent to an orphanage by her father. Louis has never forgotten Lyla and does not know about Evan. Both have since given up music careers.
Evan has a strong faith that as long as he follows the music he hears and reacts to it, he will have a chance to be found by his parents. He makes his way to New York City, where he is taken in by a man known as "Wizard" (Robin Williams), who houses various orphans and runaways, employing them to play music on the streets and taking a large cut of their tips. Evan immediately proves to be a musical child prodigy. Wizard enlists him and gives him the name "August Rush", convincing him he will be sent back to the orphanage if his real name is ever discovered.
Lyla only discovers that her son is alive when her father, knowing that he is dying, confesses what actually happened. Lyla immediately sets out to New York to look for her 11-year-old son. Meanwhile, she begins playing the cello again, having been called back to the New York Philharmonic. At about the same time, Louis reconciles with his bandmates.
After a raid by the police, Evan takes refuge in a church, where he again impresses with his natural musical talent and is enrolled at the Juilliard School as "August Rush." A work he composes is chosen to be performed by the New York Philharmonic in Central Park, but Wizard barges into a rehearsal, and Evan reluctantly follows him back to his life of performing music on the streets.
Meanwhile, Lyla has discovered Evan's identity and has decided to stay in New York while searching for her son. While there, she decides to resume her cello career. She is then chosen to play in the same concert, which features Evan's piece. Louis, being wrongly told that Lyla has since married, also returns to New York to resume playing with his former band. He has a chance meeting with Evan in Washington Square Park and they play music together, although neither knows who the other is.
The night of the concert, Evan finally chooses to run from Wizard in favor of performing at his concert. In the meantime, Louis races to the park when he sees Evan's pseudonym along with Lyla's name on a sign billing the concert. Evan conducts his piece, and at its conclusion, he turns around to see Lyla and Louis standing hand in hand, and he finally makes the connection that they are his parents
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